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Öğe Leaf Area Regulates the Growth Rates and Seed Yield of Soybean (Glycine max L. Merr.) in Intercropping System(Springer, 2022) Raza, Muhammad Ali; Gul, Hina; Hasnain, Ali; Bin Khalid, Muhammad Hayder; Hussain, Sajad; Abbas, Ghulam; Ahmed, WaqasBiotic or abiotic stresses reduce leaf area of soybean plants in the intercropping system, especially during critical reproductive growth phase (from pod-initiation to physiological-maturity) of soybean, which finally influences yield and yield components. However, total yield loss due to reduction in soybean leaf area under maize/soybean intercropping system is still unclear. In a three-year field study, an experiment consisted of four treatments: no removal of trifoliate (CK, 100% leaf area), removal of three-trifoliate (SI, 85% leaf area), removal of six-trifoliate (SII, 70% leaf area), and removal of nine-trifoliate (SIII, 55% leaf area) from the top of the soybean canopy under maize/soybean intercropping. These defoliation treatments were applied at the pod initiation (R-3) stage by removing the different number of fully developed trifoliate from the top of the soybean canopy in maize/soybean intercropping system. Results revealed that defoliation significantly decreased total dry matter accumulation and partitioning to vegetative and reproductive organs. Compared with CK (no defoliation), treatments SI, SII, and SIII reduced crop growth rate (by 25%, 46%, and 75%), reproductive growth rate (by 21%, 44%, and 63%), pod-initiation (by 11%, 23%, and 32%), while increased pod-abscission (by 11%, 20%, and 37%) and photosynthetic-rate (by 8%, 19%, and 28%), respectively at physiological-maturity. These negative responses reduced pods plant(-1) by 16%, 32%, and 49% and seeds plant(-1) by 20%, 34%, and 46% in SI, SII, and SIII, respectively, compared to non-defoliated. Overall, in SI, SII, and SIII, soybean produced 80%, 67%, and 55% of CK yield. Results implied that any change in leaf area of intercropped-soybean, especially during reproductive phase, will directly affect the availability of photoassimilates and nutrients for developing pods and seeds. Thus, more attention should be paid to improve leaf area of intercropped soybean for the high productivity of intercropping systems via appropriate variety selection or planting arrangement. Furthermore, breeders can evolve new soybean varieties, particularly for the intercropping systems, which can cope with the shading effects of tall crops in intercropping systems. Future studies are needed to understand the internal signaling and the molecular mechanism controlling in soybean in intercropping system.Öğe Maize/soybean strip intercropping produces higher crop yields and saves water under semi-arid conditions(Frontiers Media Sa, 2022) Raza, Muhammad Ali; Yasin, Hassan Shehryar; Gul, Hina; Qin, Ruijun; Mohi Ud Din, Atta; Khalid, Muhammad Hayder Bin; Hussain, SajadSustainable increases in crop production require efficient use of resources, and intercropping can improve water use efficiency and land productivity at reduced inputs. Thus, in a three-year field experiment, the performance of maize/soybean strip intercropping system differing with maize plant density (6 maize plants m-2, low, D1; 8 maize plants m-2, medium, D2; and 10 maize plants m-2, high, D3) was evaluated in comparison with sole maize or soybean cropping system. Results revealed that among all intercropping treatments, D2 had a significantly higher total leaf area index (maize LAI + soybean LAI; 8.2), total dry matter production (maize dry matter + soybean dry matter; 361.5 g plant-1), and total grain yield (maize grain yield + soybean grain yield; 10122.5 kg ha-1) than D1 and D3, and also higher than sole maize (4.8, 338.7 g plant-1, and 9553.7 kg ha-1) and sole soybean (4.6, 64.8 g plant-1, and 1559.5 kg ha-1). The intercropped maize was more efficient in utilizing the radiation and water, with a radiation use efficiency of 3.5, 5.2, and 4.3 g MJ-1 and water use efficiency of 14.3, 16.2, and 13.3 kg ha-1 mm-1, while that of intercropped soybean was 2.5, 2.1, and 1.8 g MJ-1 and 2.1, 1.9, and 1.5 kg ha-1 mm-1 in D1, D2, and D3, respectively. In intercropping, the land and water equivalent ratios ranged from 1.22 to 1.55, demonstrating that it is a sustainable strategy to improve land and water use efficiencies; this maximization is likely associated with the species complementarities for radiation, water, and land in time and space, which resulted in part from competition avoidance responses that maximize the economic profit (e. g., 1300 US $ ha-1 in D2) over sole maize (798 US $ ha-1) or sole soybean (703 US $ ha-1). Overall, these results indicate that optimizing strip intercropping systems can save 20-50% of water and land, especially under the present scenario of limited resources and climate change. However, further research is required to fully understand the resource capture mechanisms of intercrops in intercropping.Öğe Salinity Stress in Wheat (Triticum aestivum L.) in the Changing Climate: Adaptation and Management Strategies(Frontiers Media Sa, 2021) EL Sabagh, Ayman; Islam, Mohammad Sohidul; Skalicky, Milan; Raza, Muhammad Ali; Singh, Kulvir; Hossain, Mohammad Anwar; Hossain, AkbarWheat constitutes pivotal position for ensuring food and nutritional security; however, rapidly rising soil and water salinity pose a serious threat to its production globally. Salinity stress negatively affects the growth and development of wheat leading to diminished grain yield and quality. Wheat plants utilize a range of physiological biochemical and molecular mechanisms to adapt under salinity stress at the cell, tissue as well as whole plant levels to optimize the growth, and yield by off-setting the adverse effects of saline environment. Recently, various adaptation and management strategies have been developed to reduce the deleterious effects of salinity stress to maximize the production and nutritional quality of wheat. This review emphasizes and synthesizes the deleterious effects of salinity stress on wheat yield and quality along with highlighting the adaptation and mitigation strategies for sustainable wheat production to ensure food security of skyrocketing population under changing climate.Öğe Spatial differences influence nitrogen uptake, grain yield, and land-use advantage of wheat/soybean relay intercropping systems(Nature Portfolio, 2023) Raza, Muhammad Ali; Din, Atta Mohi Ud; Zhiqi, Wang; Gul, Hina; Rehman, Sana Ur; Bukhari, Birra; Haider, ImranCereal/legume intercropping is becoming a popular production strategy for higher crop yields and net profits with reduced inputs and environmental impact. However, the effects of different spatial arrangements on the growth, grain yield, nitrogen uptake, and land-use advantage of wheat/soybean relay intercropping are still unclear, particularly under arid irrigated conditions. Therefore, in a three-year field study from 2018 to 2021, soybean was relay intercropped with wheat in different crop configurations (0.9 m, narrow strips; 1.8 m, medium strips; and 2.7 m, wide strips), and the results of intercropping systems were compared with their sole systems. Results revealed that intercrops with wide strips outperformed the narrow and medium strips, when the objective was to obtain higher total leaf area, dry matter, nitrogen uptake, and grain yield on a given land area due to reduced interspecific competition between intercrops. Specifically, at maturity, wide strips increased the dry matter accumulation (37% and 58%) and its distribution in roots (37% and 55%), straw (40% and 61%), and grains (30% and 46%) of wheat and soybean, respectively, compared to narrow strips. This enhanced dry matter in wide strips improved the soybean's competitive ability (by 17%) but reduced the wheat's competitive ability (by 12%) compared with narrow strips. Noticeably, all intercropping systems accumulated a significantly higher amount of nitrogen than sole systems, revealing that wheat/soybean relay intercropping requires fewer anthropogenic inputs (nitrogen) and exerts less pressure on the ecosystem than sole systems. Overall, in wide strips, intercropped wheat and soybean achieved 62% and 71% of sole wheat and soybean yield, respectively, which increased the greater total system yield (by 19%), total land equivalent ratio (by 24%), and net profit (by 34%) of wide strips compared to narrow strips. Our study, therefore, implies that the growth parameters, grain yields, nutrient accumulation, and land-use advantage of intercrop species could be improved with the proper spatial arrangement in cereal/legume intercropping systems.